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Issue: Vol. 5332 No. 1: 18 Aug. 2023
Type: Monograph
Published: 2023-08-18
DOI: 10.11646/zootaxa.5332.1.1
Page range: 1-307
Abstract views: 992
PDF downloaded: 56

An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini

ICAR–National Research Centre for Banana; Thogamalai Road; Thayanur Post; Tiruchirappalli 620102; Tamil Nadu; India
Coleoptera Coccinelloidea pictorial guide checklist South Asia

Abstract

An illustrated guide to the tribe Coccinellini (Coleoptera: Coccinellidae: Coccinellinae) of the Indian Subcontinent comprising 119 species of 30 genera is presented. Brief diagnostic accounts of the genera and the species included therein are given with supporting illustrations of diagnostic characters, including genitalia, wherever available. Details of distribution, prey and plant associations, and seasonal occurrence are provided based on collections, label data and literature. For 31 species, illustrations of the life stages are included to facilitate the field identification of the immature stages. Adalia puetzi Iablokoff-Khnzorian, 1986 is synonymized with Adalia simmondsi Kapur & Sudha Rao, 1962 (new synonym). Oenopia smetanai Canepari, 1997 is removed from synonymy with O. flavidbrunna Jing, 1986 and restored as a valid species (status revised). Illeis kapuri Anand, Gupta & Ghai is transferred to Ortalia Mulsant, 1850 (new combination). Two new species, Harmonia andamanensis sp. n. and Phrynocaria prathapani sp. n. are described and illustrated from the Andamans and the Western Ghats, respectively. Coelophora lushuiensis (Jing, 1992) is documented as a new record for India (Manipur). An updated checklist of the Coccinellini of the Indian Subcontinent is also provided.

 

References

  1. Abdel-Dayem, M.S., Fad, H.H., El-Torkey, A.M., Elgharbawy, A.A., Aldryhim, Y.N., Kondratieff, B.C., Al Ansi, A.N. & Aldhafer, H.M. (2017) The beetle fauna (Insecta, Coleoptera) of the Rawdhat Khorim National Park, Central Saudi Arabia. ZooKeys, 653, 1–78. https://doi.org/10.3897/zookeys.653.10252
  2. Abdolahi, R., Nedvěd, O. & J. Nozari, J. (2018) New data on some Coccinellidae (Coleoptera) from Iran. Acta Phytopathologica et Entomologica Hungarica, 53 (1), 83−90. https://doi.org/10.1556/038.52.2017.035
  3. Afroze, S. (2000) Bioecology of the coccinellid Anegleis cardoni (Weise) (Coleoptera: Coccinellidae), an important predator of aphids, coccids and pseudococcids. Journal of Entomological Research, 24, 55–62.
  4. Afroze, S. & Shafee, A.A. (1998) A new species of the genus Micraspis (Coccinellidae: Coleoptera) from India. Columban Journal of Life Sciences, 6 (2), 324–327.
  5. Afroze, S. & Shujauddin (1998) Bioecology of Synia melanaria (Coccinellidae: Coleoptera) predating on Coptosoma ostensum Distant. Journal of Entomological Research, 22, 329–336.
  6. Afsana, N. & Islam, Z. (2001) Abundance of ladybird beetles in irrigated rice environment in Bangladesh, diurnal activity pattern of Micraspis discolor (Fab.) and its influence on rice grain sterility. Bangladesh Journal of Entomology, 11, 55–66.
  7. Agarwala, B.K. & Ghosh, A.K. (1988) Prey records of aphidophagous Coccinellidae in India: A review and bibliography. Tropical Pest Management, 34, 1–14. https://doi.org/10.1080/09670878809371196
  8. Agarwala, B.K. & Yasuda, H. (2000) Competitive ability of ladybird predators of aphids: A review of Cheilomenes sexmaculata (Fabr.) (Coleoptera: Coccinellidae) with a worldwide checklist of preys. Journal of Aphidology, 14, 1–20.
  9. Agarwala, B.K., Saha, S. & Ghosh, A.K. (1984) Studies on Ceratovacuna silvestrii (Takahashi) (Hemiptera: Aphididae) and its predator Anisolemnia dilatata (Fab.) on Bambusa arundinacea. Records of the Zoological Survey of India, 81, 23–42. https://doi.org/10.26515/rzsi/v81/i3-4/1984/161216
  10. Ali, M., Naqvi, A.N., Perveen, R., Ahmad, K. & Hussain, I. (2014) First record of the tribe Bulaeini (Coleoptera Coccinellidae) with generic and species descriptions from Pakistan. Pakistan Journal of Zoology, 46 (5), 1475–1478.
  11. Amman G.D. & Speers C.F. (1964) Release of predators of the balsam woolly aphid in North Carolina. United States Forest Service Southeast Forest Experiment Station Research Notes, 153, 1–4.
  12. Anand, R.K., Gupta, A.K. & Ghai, S. (1990) A new species of Illeis Muls. from North India (Insecta, Coleoptera: Coccinellidae). Reichenbachia, 27 (2), 131–134.
  13. Aruggoda, A.G.B., Ren, S. & Baoli, Q. (2010) Molecular phylogeny of ladybird beetles (Coccinellidae: Coleoptera) inferred from mitochondria 16S rDNA sequences. Tropical Agricultural Research, 21, 209–217. https://doi.org/10.4038/tar.v21i2.2602
  14. Ashfaque, M., Ullah, F. & Rafi, M.A. (2013) Genus Coccinella (Coccinellidae: Coleoptera) from Gilgit Baltistan with two new records from Pakistan. Sarhad Journal of Agriculture, 29 (2), 239–247.
  15. Assour, H.R. & Behm, J.E. (2019) First occurrence of Cheilomenes sexmaculata (Coleoptera: Coccinellidae) on the Caribbean Island of Curaçao. Neotropical Entomology, 48, 863–865. https://doi.org/10.1007/s13744-019-00699-0
  16. Barcos, A.K., Velasco, L.I. & Medina, C. (2014) Foraging behavior of the yellow netted ladybeetle Heteroneda billardieri Crotch (Coleoptera: Coccinellidae). Philippine Agricultural Scientist, 97 (2), 155–160.
  17. Barovsky, V. (1928) De genere Ithone Sols. (Coleoptera: Coccinellidae). Revue Russe d’Entomologie, Leningrad, 22, 231–232.
  18. Beeson, C.F.C. (1941) The Ecology and Control of the Forest Insects of India and the Neighbouring Countries. Bishen Singh–Mahendrapal Singh, Dehradun, 1007 pp.
  19. Behura, B.K. (1963a) Aphids of India: A survey of published information. In: Proceedings of the First Summer School of Zoology (Simla, 1961). Government of India publications, New Delhi, pp. 25–78.
  20. Behura, B.K. (1963b) Supplement to “Aphids of India—A survey of published information”. Journal of the Utkal University (Science), Bhubaneswar, 3 (1 & 2), 40–65.
  21. Belicek, J. (1976) Coccinellidae of Western Canada and Alaska with analyses of the transmontane zoogeographic relationships between the fauna of British Columbia and Alberta (Insecta: Coleoptera: Coccinellidae). Quaestiones Entomologicae, 12 (4), 283–409.
  22. Bhattacharjee, S.S., Chakraborty, N., Kumar C.A. & Sahakundu, A.K. (1994) Control of white powdery mildew, Phyllactinia corylea (Pers.) Karst. with the lady bird beetle, Illeis indica Timb. (Coleoptera: Coccinellidae). Sericologia, 34, 485–491.
  23. Bielawski, R. (1957) Coccinellidae (Coleoptera) von Ceylon. Verhandlungen der Naturforschenden Gesellschaft in Basel, 68, 72–96.
  24. Bielawski, R. (1959) Coccinellidae (Coleopt.) von Sumba, Sumbawa, Flores, Timor und Bali. Wissenschaftliche Ergebnisse der Sumba Expedition des Museums für Völkerkunde und des Naturhistorischen Museums in Basel, 1949, 144–166.
  25. Bielawski, R. (1960) Materialien zur Kenntnis der Coccinellidae (Coleoptera). Annales Zoologici, 18, 435–458.
  26. Bielawski, R. (1961a) Bemerkung über die männlichen Genitalien von Arten de Gattung Illeis Muls., nebst Beschreibung einer neuen Art und einer Unterart (Coleoptera, Coccinellidae). Annales Zoologici, 19, 353–368.
  27. Bielawski, R. (1961b) Materialien zur kenntnis der Coccinellidae (Coleoptera). II. Annales Zoologici, 19, 385–415.
  28. Bielawski, R. (1963a) Beiträge zur Kenntnis der Coccinelliden von Afghanistan. III. Lunds Universitets Årsskrift, Neue Folge, 59 (4), 1–21.
  29. Bielawski, R. (1963b) Monographie der Epilachninae (Coleoptera, Coccinellidae) der australischen Region. Annales Zoologici, 21, 295–461.
  30. Bielawski, R. (1964a) A review of the New Guinean species of the genus Harmonia Muls. (Coleoptera: Coccinellidae). Pacific Insects, 6, 5–13.
  31. Bielawski, R. (1964b) Die Arten der Gattung Gyrocaria Timberlake (Coleoptera: Coccinellidae). Bulletin de l’Académie Polonaise des Sciences, Cl. II, Serie des Sciences Biologiques, 12, 83–90.
  32. Bielawski, R. (1964c) Die Arten der Gattung Synia Mulsant (Coleoptera: Coccinellidae). Bulletin de l’Académie Polonaise des Sciences, Cl.II, Serie des Sciences Biologiques, 12, 23–27.
  33. Bielawski, R. (1968) [new species]. In: Bielawski, R. & Chûjô, M., Coleoptera from Southeast Asia arranged by M. Chûjô (VII). Coccinellid-beetle from E. Nepal, Thailand, Cambodia, Viet-Nam, S. Korea and Is. Ambon, with descriptions of 4 new species and 33 figures. Transactions of the Shikoku Entomological Society, 9, pp. 119–128.
  34. Bielawski, R. (1971) Über Coccinellidae (Coleoptera) aus Nepal. Khumbu Himal, Innsbruck–München. Ergebnisse Forsch Unternehemens Nepal Himalaya, 4, 1–9.
  35. Bielawski, R. (1972) Die Marienkäfer (Coleoptera: Coccinellidae) aus Nepal. Fragmenta Faunistica, 18, 283–312. https://doi.org/10.3161/00159301FF1972.18.16.283
  36. Bielawski, R. (1979) Ergebnisse der Bhutan–Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Coccinellidae. Entomologica Basiliensia, 4, 83–125.
  37. Bielawski, R. & Chujo, M. (1964) Coleoptera from Southeast Asia (III). 21. Family Coccinellidae. Nature and Life in South east Asia, 3, 226–228.
  38. Bielawski, R. & Fürsch, H. (1960) Zwei neue Coccinelliden aus Burma. Mitteilungen der Münchner Entomologischen Gesellschaft, 50, 68–71.
  39. Biranvand, A., Nedvěd, O., Nattier, R., Nepaeva, E. & Haelewaters, D. (2021) Review of the genus Hippodamia (Coleoptera: Coccinellidae) in the Palearctic region. Oriental Insects, 55, 2, 293–304. https://doi.org/10.1080/00305316.2020.1763871
  40. Blackburn, T. (1892) Further notes on Australian Coleoptera, with descriptions of new genera and species. XII. Transactions of the Royal Society of South Australia, 15 (2), 207–261.
  41. Boisduval, J.B.A. (1835) Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, Pendant les Années 1826–1827–1828–1829, sous le Commandement de M. J. Dumont d’Urville. Faune Entomologique de l’Océan Pacifique, avec l’illustration des Insectes Nouveaux Recueillis Pendant le voyage. Deuxième Partie. Coléoptères et autres ordres. J. Tastu, Paris, 716 pp. https://doi.org/10.5962/bhl.title.35819
  42. Boopathi, T., Singh, S.B., Dutta, S.K., Dayal, V., Singh, A.R., Chowdhury, S., Ramakrishna, Y., Aravintharaj, R., Shakuntala, I. & Lalhruaipuii, K. (2020) Harmonia sedecimnotata (F.): Predatory potential, biology, life table, molecular characterization, and field evaluation against Aphis gossypii Glover. Scientific Reports, 10, 3079. https://doi.org/10.1038/s41598-020-59809-3
  43. Booth, R.G. (1993) Identification Services Report. List No. 11773 Asia. International Institute of Entomology, London. [unknown pagination]
  44. Booth, R.G. (1997) A review of the species of Calvia (Coleoptera: Coccinellidae) from the Indian subcontinent, with descriptions of two new species. Journal of Natural History, 31, 917–934. https://doi.org/10.1080/00222939700770451
  45. Booth, R.G. & Pope, R.D. (1989) A review of the type material of Coccinellidae (Coleoptera) described by F.W. Pope and by E. Mulsant in the Hope Entomological Collections, Oxford. Entomologica Scandinavica, 20 (3), 343–370. https://doi.org/10.1163/187631289X00366
  46. Booth, R.G., Cox, M.L. & Madge, R.B. (1990) IIE Guides to Insects of Importance to Man. 3. Coleoptera. CAB International, Wallingford, 384 pp.
  47. CABI (2002) Crop Protection Compendium. Global Module. 2nd Edition. CAB International, Wallingford. [CD-ROM]
  48. Canepari, C. (1986) Su alcuni Coccinellidi dell’India e Nepal settentrionale del Museo di Storia Naturale di Ginevra (Coleoptera: Coccinellidae). Revue Suisse de Zoologie, 93, 21–36. https://doi.org/10.5962/bhl.part.79677
  49. Canepari, C. (1990) Who is the author of several genera of Coccinellidae: Chevrolat or Dejean? Coccinella, 2 (1), 38–39.
  50. Canepari, C. (1997) Coccinellidae (Coleoptera) from the Nepal Himalayas. Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 565 (65), 1–65.
  51. Canepari, C. (2003). Coccinellidae (Insecta: Coleoptera) of Nepal from the collection of the Naturkundemuseum Erfurt. In: Biodiversity and natural heritage of the Himalaya (Hartmann, M. & Baumbach, H. (Eds.), Verein der Freunde & Förderer des Naturkundemuseums Erfurt e.V. Naturkundemuseum, Erfurt, pp. 261–265.
  52. Canepari, C. (2012) Coccinellidae (Insecta: Coleoptera) of Nepal Himalaya in the Naturkundemuseum Erfurt. In: Hartmann, M. & Weipert, J. (Eds.), Biodiversität und Naturausstattung im Himalaya IV. Verein der Freunde und Forderer des Naturkundemuseums Erfurt e.V., Erfurt, pp. 357–390.
  53. Cao, C. & Pu, T. (1985) A new species and new record of Coccinellidae from Guangxi Zhuangzu autonomous region. Entomotaxonomia, 7, 273–275.
  54. Ceryngier, P. & Hodek, I. (1996) Enemies of Coccinellidae. In: Hodek, I. & Honěk, A. (Eds.), Ecology of Coccinellidae. Kluwer Academic Publishers, Dordrecht, pp. 319–350. https://doi.org/10.1007/978-94-017-1349-8_8
  55. Ceryngier, P., Roy, H.E. & Poland, R.L. (2012) Natural enemies of ladybird beetles. In: Hodek, I. & Honěk, A. (Eds.), Ecology and behaviour of the ladybird beetles (Coccinellidae). 1st Edition. Blackwell Publishing Ltd., Chichester, pp. 375–443. https://doi.org/10.1002/9781118223208.ch8
  56. Chakrabarti, S., Ghosh, D. & Debnath, N. (1988) Developmental rate and larval voracity in Harmonia (Leis) dimidiata (Col., Coccinellidae), a predator of Eriosoma lanigerum (Hom., Aphididae) in Western Himalaya. Acta Entomologica Bohemoslovaca, 85, 335–339.
  57. Chakrabarti, S., Debnath, N. & Ghosh, D. (1995) Bioecology of Harmonia eucharis (Mulsant) (Coleoptera: Coccinellidae), an aphidophagous predator in Western Himalayas. Entomon, 20, 191–196.
  58. Channabasavanna, G.P. & Puttarudriah, M. (1957) Some predators of mites in Mysore. The Mysore Agricultural Journal, 32 (3–4), 179–185.
  59. Chapin, E.A. (1965) Coleoptera, Coccinellidae. Insects of Micronesia, 16, 189–254.
  60. Chapuis, F. (1876) Famille LXXII, Coccinellides. In: Lacordaire, T. & Chapuis, F. (Eds.), Histoire Naturelle des Insectes, Genera des Coleopteres. Vol. 12. Roret, Paris, pp. 149–259.
  61. Chatterjee, N.C. & Bose, M. (1933) Entomological investigations on the spike disease of sandal (17). Coccinellidae (Col.). Supplementary data. Indian Forest Records, 19 (7), 1–10.
  62. Che, L.H., Zhang, P., Deng, S.H., Escalona, H.E., Wang, X., Li, Y., Pang, H., Vandenberg, N., Ślipiński, A., Tomaszewska, W. & Liang, D. (2021) New insights into the phylogeny and evolution of lady beetles (Coleoptera: Coccinellidae) by extensive sampling of genes and species. Molecular Phylogenetics and Evolution, 156, 107045. https://doi.org/10.1016/j.ympev.2020.107045
  63. Chengyi, C. (Ed.) (1992) Fauna of Yunnan. Coccinellidae. Yunnan Science & Technology Pub., Kunming, 243 pp.
  64. Chunram, S. & Sasaji, H. (1980) A contribution to the Coccinellidae (Coleoptera) of Thailand. Oriental Insects, 14 (4), 473–491. https://doi.org/10.1080/00305316.1980.10434833
  65. CIBC (1984) Survey for natural enemies of diaspine scale insects in South India, US PL-480 project. Final Technical Report (unpublished). Commonwealth Institute of Biological Control, Indian station, Bangalore, 87 pp.
  66. Clausen, C.P. (1940) Entomophagous Insects. McGraw-Hill Book Co., Inc., New York, 688 pp., 257 figs.
  67. Clausen, C.P. (Ed.) (1978) Introduced parasites and predators of arthropod pests and weeds: a world review. Agriculture Handbook No. 480. U.S. Department of Agriculture, Washington, D.C., 545 pp.
  68. Coutanceau, J. (2008) Le genre Harmonia (Mulsant, 1846) (Coleoptera Coccinellidae). Harmonia, 1, 4–16. [in French]
  69. Crotch, G.R. (1871) List of Coccinellidae. Printed by the author, Cambridge, 8 pp.
  70. Crotch, G.R. (1874) A Revision of the Coleopterous Family Coccinellidae. Janson, London, 311 pp. https://doi.org/10.5962/bhl.title.8975
  71. Crowson, R.A. (1955) The natural classification of the families of Coleoptera. Nathaniel Lloyd, London, 187 pp.
  72. Das, P., Saha, G.K., Gupta, D. & Chandra, K. (2020a) New faunistic record of two species of genus Calvia (Coleoptera: Coccinellidae: Coccinellini) from India and Myanmar. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”, 63 (2), 195–201. https://doi.org/10.3897/travaux.63.e57502
  73. Das, P., Chandra, K. & Gupta, D. (2020b) The ladybird beetles (Coleoptera: Coccinellidae) of Arunachal Pradesh, East Himalaya, India with new combinations and new country records. Bonn Zoological Bulletin, 69 (1), 27–44.
  74. Das, P., Saha, G.K., Chandra, K. & Gupta, D. (2022) New to the fauna of India ladybird beetles (Coleoptera: Coccinellidae). Far Eastern Entomologist, 449, 9–17. https://doi.org/10.25221/fee.449.2
  75. Dauguet, P. (1949) Les Coccinellini de France. Editions de l’Entomologiste, Paris, 46 pp.
  76. Debaraj, Y. & Singh, T.K. (2000) Field recognition of the different developmental stages of an aphidophagous predator, Coccinella transversalis. Annals of Plant Protection Sciences, 8, 242–244.
  77. De Geer, C. (1778) Mémoires pour servir à l’histoire des Insectes. Tome Septième. Ouvrage posthume. Pierre Hesselberg, Stockholm, xii + 950 pp., 49 pls.
  78. Dejean, P.F.M.A. (1836–37) Catalogues des Coléoptères de la collection de M. le Comte Dejean. Troisièmeme Edition. Revue, Corrigée et Augmentée. Mequignon-Marvis père et fils, Paris, xiv + 503 pp. [Fascicle 5, pp. i–xiv + 385–503 (May 1837)] https://doi.org/10.5962/t.173109
  79. Dejean, A., Orivel, J. & Gibernau, M. (2002) Specialized predation on plataspid heteropterans in a coccinellid beetle: adaptive behavior and responses of prey attended or not by ants. Behavioral Ecology, 13, 154–159. https://doi.org/10.1093/beheco/13.2.154
  80. Deng, G.R., Yang, H.H. & Jin, M.X. (1987) Augmentation of coccinellid beetles for controlling sugarcane woolly aphid. Chinese Journal of Biological Control, 3, 166–168.
  81. Devi, S.S. (1989) Studies on taxonomy and biology of aphidophagous coccinellids (Coleoptera: Coccinellidae) of Manipur and Nagaland. Ph.D. dissertation submitted to Manipur University, Canchipur, 233 pp.
  82. Dobzhansky, T. (1925) Die palaarktischen Arten der Gattung Coccinula Dobzhansky. Zoologischer Anziger, 64, 277–284.
  83. Dobzhansky, T. (1926) Die Palaarktischen Arten der Gattung Coccinella L. Revue Russe d’Entomologie, 20, 16–32.
  84. Dobzhansky, T. (1927) Neue und wenig bekannte Coccinelliden. Revue Russe d’Entomologie, 21, 212–217.
  85. Dohrn, C.A. (1882) Exotisches. Stettiner Entomologische Zeitung, 43, 363–374.
  86. Dorji, C., Loday, P. & Vorst, O. (2017) A field guide to the common lady beetles of Bhutan. National Biodiversity Centre, Serbithang, Thimphu, 53 pp.
  87. Dorji, C., Loday, P. & Vorst, O. (2019) A preliminary checklist of the Coccinellidae of Bhutan (Insecta: Coleoptera). Zootaxa, 4712 (4), 497–530. https://doi.org/10.11646/zootaxa.4712.4.2
  88. Dorji, C., Lhamo, T. & Wangchuk, J. (2021) Six species of Coccinellidae new to Bhutan (Insecta: Coleoptera). Oriental Insects, 56 (2), 187–197. https://doi.org/10.1080/00305316.2021.1930604
  89. Duang, T.H. & Frederick, W.S. (1974) Morphology and taxonomy of the known pupae of Coccinellidae (Coleoptera) of North America, with a discussion of phylogenetic relationships. Contributions of the American Entomological Institute, 10 (6), 1–125.
  90. Easwaramoorthy, S., Nirmala, R. & Santhalakshmi, G. (2001) Biology and predatory potential of Micraspis univittata (Hope), a coccinellid predator recorded in sugarcane ecosystem. Journal of Biological Control, 15, 97–100.
  91. Emden, F. Van (1949) The larvae of British beetles: VII. Coccinellidae. Entomologists Monthly Magazine, 85, 265–283.
  92. Escalona, H.E., Zwick, A., Li, H.S., Wang, X., Pang, H., Hartley, D., Jermiin, L.S., Nedvěd, O., Misof, B., Niehuis, O., Ślipiński, A. & Tomaszewska, W. (2017) Molecular phylogeny reveals food plasticity in the evolution of true ladybird beetles (Coleoptera: Coccinellidae: Coccinellini). BMC Evolutionary Biology, 17, 151. https://doi.org/10.1186/s12862-017-1002-3
  93. Fabricius, J.C. (1775) Systema Entomologiae, sistens Insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensburgi et Lipsiae, 832 pp. https://doi.org/10.5962/bhl.title.36510
  94. Fabricius, J.C. (1781) Species Insectorum exhibitiens eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Vol. 1. Impensis Carol. Ernest, Bohnii, Hamburgi et Kiloonii, 382 pp. https://doi.org/10.5962/bhl.title.36509
  95. Fabricius, J.C. (1787) Mantissa Insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Vol. 1. Impensis Christ. Gottl. Proft, Hafniae, XX + 348 pp. https://doi.org/10.5962/bhl.title.11657
  96. Fabricius, J.C. (1792) Entomologia Systematica Emendata et Aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Tom I. Pars I. Impensis Christ. Gottl. Proft., Hafniae, 535 pp. https://doi.org/10.5962/bhl.title.122153
  97. Fabricius, J.C. (1798) Supplementum entomologiciae systematicae. impensis Christ. Gottl. Proft, Hafniae, 53 pp.
  98. Fabricius, J.C. (1801) Systema Eleutheratorum secundum ordines, genera species adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Impensis Bibliopolii Academici Novi, Kiliae, 506 pp.
  99. Faldermann, F. (1835) Coleopterorum ab ill. Bungio in China boreali, Mongolia et montibus Altaicis collectorum, nee non ab ill. Turczaninoffio et Stschukino e provincia Irkutzk missorum illustrationes. Memoires de l’Academie Imperiale des Sciences des Saint Petersbourg, 2, 337–464.
  100. Fürsch, H. (1960) Neue Coccinellidae aus dem Museum Frey. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei München, Bd. 11 (1), 298–303.
  101. Fürsch, H. (1964) Neue Gesichtspunkte zur Beurteilung des Gattungs namens Micraspis Dejean (Col., Coccinellidae). Nachrichtenblatt der Bayerischen Entomologen, 13, 70–72.
  102. Fürsch, H. (1967) Familie: Coccinellidae (Marienkäfer). In: Freude, H., Harde, K.W. & Lohse, G.A. (Eds.), Die Käfer Mitteleuropas. Band 7. Clavicornia. Goecke und Evers, Krefeld, pp. 227–278.
  103. Fürsch, H. (1981) Coccinella marussii Kapur, eine neue Art in der Turkei (Coleoptera: Coccinellidae). Nachrichtenblatt der Bayerischen Entomologen, 30 (5), 81–84.
  104. Fürsch, H. (1987) Die Afrikanischen Vertreter der Gattungen Micraspis, Declivitata und Xanthadalia (Coleoptera: Coccinellidae). Mitteilungen Muenchener Entomologischen Gesellschaft, 77, 5–31.
  105. Fürsch, H. (1989) Discussion about Iablokoff-Khnzorian’s paper of 1986 about Old World Coccinellidae. Coccinella, 2, 28–33.
  106. Fürsch, H. (1996) Taxonomy of Coccinellids. Coccinella, 6, 28–30.
  107. Fürsch, H. (2005) Bulaea MULSANT, 1850 und Isora MULSANT, 1850 sind synonym. Entomologische Zeitschrift Stuttgart, 115, 227–232.
  108. Ganglbauer, L. (1899) Die Kafer von Mitteleuropa. 3. Familienreihe Staphylinoidea, 2. Theil: Familienreihe Clavicornia. 33. Family Coccinellidae. Gerold, Wien, 1046 pp.
  109. Gebler, F. (1832) Notice sur les Colèoptéres qui se trouvent dans le district des mines de Nertchinsk, dans la Sibèrie orientale, avec la description de quelques espëces nouvelles. Nouveax Mèmoires de la Sociètè Impèriale des Naturalistes de Moscou, 2, 23–78.
  110. Ghani, M.A. (1962) A note on the identity of some species of the genus Ballia (Coleoptera: Coccinellidae). Proceedings of the Royal Entomological Society of London (B), 31, 7–8. https://doi.org/10.1111/j.1365-3113.1962.tb01193.x
  111. Ghani, M.A. & Muzaffar, N. (1974) Relations between the parasite-predator complex and the host–plants of scale insects in Pakistan. Miscellaneous publication No. 5. Commonwealth Institute of Biological Control, CAB, Slough, 92 pp.
  112. Ghorpade, K.D. (1976) An undescribed species of Illeis (Coleoptera: Coccinellidae) from South India. Oriental Insects, 10, 579–585. https://doi.org/10.1080/00305316.1976.10434528
  113. Ghorpade, K.D. (1979a) On some Coccinellidae (Coleoptera) attracted to light in India. Current Research, 8, 114.
  114. Ghorpade, K.D. (1979b) Ballia eucharis (Coleoptera: Coccinellidae) breeding on Cicadellidae (Homoptera) at Shillong. Current Research, 8, 113.
  115. Ghosh, A.K., Lahiri, A.R. & Biswas, S. (1976) Preliminary observation on the development of a predaceous lady-bird beetle Ballia eucharis Mulsant (Coleoptera: Coccinellidae). Newsletter, Zoological Survey of India, 2, 129–130.
  116. Ghosh, A.K., Biswas, S. & Lahiri, A.R. (1977) A preliminary study of the insect fauna of Meghalaya. 7. Coleoptera: Coccinellidae. Bulletin of the Meghalaya Science Society, 2, 11–15.
  117. Ghosh, D., Debnath, N. & Chakrabarti, S. (1986) Development of three species of Oenopia (Coleoptera: Coccinellidae) predating on aphids of Garhwal Himalayas. In: Proceedings of the 3rd Oriental Entomology Symposium, 21–24 February1984. Vol. 1. Association for the Advancement of Entomology, University of Kerala, Kariavattom, pp. 173–176.
  118. Goeze, J.A.E. (1777) Entomologische Beytraege zu des Ritters Linné zwölften Ausgabe des Natursystems. Erster Theil. Weidmanns Erben und Reich, Leipzig, xiv + 736 pp. https://doi.org/10.5962/bhl.title.45974
  119. Gordon, R.D. (1985) The Coccinellidae of America, north of Mexico. Journal of the New York Entomological Society, 93, 1–912.
  120. Gordon, R.D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1–46.
  121. Gordon, R.D. & Vandenberg, N. (1995) Larval systematics of North American Coccinella L. (Coleoptera: Coccinellidae). Entomologica Scandinavica, 26, 67–86.
  122. Gorham, H.S. (1894a) On the Coccinellidae from India in the collection of Mr. H.E. Andrewes of the Indian Forestry Service. Annales de la Société Entomologique de Belgique, 38, 200–208.
  123. Gorham, H.S. (1894b) Insectes du Bengale. Observations on some species of the family Coccinellidae, collected near Konbir and Mandar, India, Bengal, by P. Cardon. Annales de la Société Entomologique de Belgique, 38, 209–211.
  124. Gorham, H.S. (1895) On the Coccinellidae collected by Mr. L. Fea in Burma. Annali del Museo Civico di Storia Naturale, Genova, Series 2, 34, 683–695. https://doi.org/10.5962/bhl.part.16769
  125. Gorham, H.S. (1903) On Coleoptera collected in India by MM. H.E. and H. L. Andrewes, Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae. Annales de la Société Entomologique de Belgique, 47, 323–347.
  126. Guérin-Méneville, F.E. (1835) Iconographie du Regne Animale de Cuvier. Vol. 2. Insectes. J. B. Balliere, Libraire de l’Académie Royale de Médicine, Paris, 104 pls. [pls. 1829–1837, text 1844]
  127. Gurung, B., Ponnusamy, N. & Pal, S. (2019) Species diversity of predaceous coccinellids in different crop ecosystems under the hilly and terai region of West Bengal (India). Ecology, Environment and Conservation, 25 (2), 636–642.
  128. Hayat, A., Khan, M. R. & Naz, F. (2017) Subfamilies Coccinellinae and Coccidullinae (Coccinellidae: Coleoptera) with New Records from AJK, Pakistan. Journal of Applied Environmental and Biological Sciences, 7 (4), 12–66.
  129. Hoang, D.N. (1982) A new genus and some new species of subfamily Coccinellinae, (Coleoptera: Coccinellidae) from Vietnam. Tap chi Sinh Vat Hoc, 4 (3), 10–15.
  130. Hoang, D.N. (1983a) A new genus of ladybird beetles (Coleoptera, Coccinellidae) from Vietnam. In: Medvedev, L.N. (Ed.), Fauna and ecology of the animals of Vietnam. Nauka, Moscow, pp.150–156. [in Russian]
  131. Hoang, D.N. (1983b) Coccinellidae of Vietnam. Part. 2. Nha zuat ban khoa hoc va ky thuat, Hanoi, 159 pp. [in Vietnamese with English summary]
  132. Hoang, D.N. (1984) Three new species of the coccinellid genus Nedina Hoang (Coleoptera, Coccinellidae) from South Viet Nam. Entomologicheskoe Obozrenie, 63 (1), 68–73. [in Russian]
  133. Hodek, I. & Honěk, A. (1996) Ecology of Coccinellidae. Kluwer Academic Publishers, Dordrecht, xvi + 464 pp. https://doi.org/10.1007/978-94-017-1349-8
  134. Hodek, I., van Emden, H.F. & Honěk, A. (2012) Ecology and Behaviour of the Ladybird Beetles (Coccinellidae). Blackwell Publishing Ltd., Chichester, 561 pp. https://doi.org/10.1002/9781118223208
  135. Hope, F.W. (1831) s.n. In: Gray, J.E., Synopsis of the new species of Nepaul insects in the collection of Major General Hardwicke. The Zoological Miscellany, London, pp. 21–32.
  136. Hope, F.W. (1840) The Coleopterist’s Manual, Part the Third, Containing Various Families, Genera, and species of Beetles Recorded by Linnaeus and Fabricius. Also Descriptions of Newly Discovered and Unpublished Insects, London, 191 pp.
  137. Hope, F.W. (1843) Descriptions of the coleopterous insects sent to England by Dr. Cantor from Chusan and Canton, with observations on the entomology of China. Annals and Magazine of Natural History, 11, 62–66.
  138. Hummel, A.D. (1827) Essais Entomologiques. No. VI. Insectes de 1826. s.n., St. Petersbourg. 50 pp.
  139. Iablokoff-Khnzorian, S.M. (1972) Les types de Coccinellidae de la collection Motschulsky (Coléoptères Coccinellidae). Nouvelle Revue d’Entomologie, 2 (2), 163–184.
  140. Iablokoff-Khnzorian, S.M. (1976) A new species of Coleoptera Coccinellidae from China (Coleoptera, Coccinellidae). Doklady Academy of Sciences of the Armenian SSR, 63 (4), 250–255.
  141. Iablokoff-Khnzorian, S.M. (1979) Genera der Paläarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blätter für Biologie und Systematik der Käfer, 75 (1–2), 37–75.
  142. Iablokoff-Khnzorian, S.M. (1982) Les Coccinelles Coléoptères-Coccinellidae. Tribu Coccinellini des régions Paléarctique et Orientale. Boubée, Paris, 568 pp. [in French]
  143. Iablokoff-Khnzorian, S.M. (1984) Notes sur la tribu des Coccinellini (Coleoptera: Coccinellidae). Nouvelle Revue d’Entomologie, New Series, 1 (2), 203–220.
  144. Iablokoff-Khnzorian, S.M. (1986) Adalia puetzi n. sp., nouvelle espece du Pakistan (Col. Coccinellidae). Nouvelle Revue d’Entomologie, New Series, 3 (1), 80.
  145. Illahi, I., Khan, M.A., Zeya, S.B., Aslam, M. & Mittal, V. (2011) Feeding potential of a mycophagous beetle Halyzia tschitscherini (Coleoptera: Coccinellidae) against powdery mildew Phyllactinia corylea in mulberry. Indian Phytopathology, 64 (2), 140–143.
  146. Iperti, G. (1999) Biodiversity of predaceous Coccinellidae in relation to bioindication and economic importance. Agriculture, Ecosystems and Environment, 74, 323–342. https://doi.org/10.1016/S0167-8809(99)00041-9
  147. Iqbal, Z., Nasir, M.F., Bodlah, I., Qureshi, R. & Aihetasham, A. (2017) Notes on three morphs of Bulaea lichatschovii (Hummel) (Coleoptera: Coccinellidae) from Northern Pakistan. Punjab University Journal of Zoology, 32 (2), 203–208.
  148. Iqbal, Z., Nasir, M.F., Bodlah, I., Szawaryn, K., Khormizi, M.Z. & Hassan, M.A. (2019) A review of the genus Hyperaspis Chevrolat, 1837 (Coleoptera: Coccinellidae) from Pakistan. Oriental Insects, 53, 58–76. https://doi.org/10.1080/00305316.2018.1440258
  149. Irshad, M. (2001) Distribution, hosts, ecology and biotic potentials of coccinellids of Pakistan. Pakistan Journal of Biological Sciences, 4 (10), 1259–1263.
  150. Jacobson, G.G. (1916) s.n. In: Die Käfer Russlands und Westeuropas. Ein Handbuch zum Bestimmen der Käfer, Petrograd, pp. 865–1024
  151. Jadwiszczak, A. (1990) On the systematic position of the genus Subepilachna Bielawski (Coleoptera: Coccinellidae). Coccinella, 2 (2), 56–57.
  152. Jadwiszczak, A. & Pokojowczyk, A. (1990) Paramulsantina Hoang (Coleoptera: Coccinellidae), a new synonym of the genus Oenopia Mulsant. Coccinella, 2 (2), 54–55.
  153. Jing, X.L. (1986) Three new species of Oenopia Mulsant (Coccinellidae: Coleoptera). Acta Entomologica Sinica, 29 (2), 203–207.
  154. Jing, X.L. (1987) A new species of Halyzia Mulsant (Coleoptera: Coccinellidae). Acta Entomologica Sinica, 30 (2), 201–202.
  155. Jing, X.L. (1992) Coleoptera: Coccinellidae. In: Chen, S. (Ed.), Insects of the Hengduan Mountains region (I). Science Press, Beijing, pp. 541–574.
  156. Joshi, P.C., Khamashon, L. & Kaushal, B.R. (2012) Predation of Adalia tetraspilota (Hope) (Coleoptera: Coccinellidae) on green peach aphid, Myzus persicae Sulzer. Entomology, Ornithology and Herpetology, 1 (1), 1–2. https://doi.org/10.4172/2161-0983.1000101
  157. Joshi, S., Ballal, C.R. & Rao, N.S. (1999) Biotic potential of three coccinellid predators on six different aphid hosts. Journal of Entomological Research, 23, 1–7.
  158. Kamiya, H. (1965a) Comparative morphology of larvae of the Japanese Coccinellidae, with special reference to the tribal phylogeny of the family (Coleoptera). Memoirs of the Faculty of Liberal Arts, Fukui University, Series II, Natural Science, 14 (5), 83–100.
  159. Kamiya, H. (1965b) A revision of the tribe Coccinellini of Japan and the Ryukyus (Coleoptera: Coccinellidae). Memoirs of the Faculty of Liberal Arts, Fukui University, Series II, Natural Sciences, 15, 27–71.
  160. Kamoi, M. & Imata, K. (1978) Notes on the life history of Callicaria superba Mulsant (Coleoptera: Coccinellidae). Kontyu, 46, 253–256.
  161. Kapur, A.P. (1939) Bionomics of two coccinellids Adonia variegata Goeze and Brumus suturalis F. predating on aphids. Proceedings, 26th Indian Science Congress, Lahore, Pt. 3 (Abstracts), 146–147.
  162. Kapur, A.P. (1940) A note on the ladybird beetles (Coccinellidae) predating upon the cane whitefly, Aleurolobus barodensis Maskell. Current Science, 9, 134.
  163. Kapur, A.P. (1942) Bionomics of some Coccinellidae predaceous on aphids and coccids in north India. Indian Journal of Entomology, 4, 49–66.
  164. Kapur, A.P. (1943) On the biology and structure of the coccinellid, Thea bisoctonotata Muls. in north India. Indian Journal of Entomology, 5, 165–171.
  165. Kapur, A.P. (1948) A revision of the tribe Aspidimerini Weise (Coleoptera: Coccinellidae). Transactions of the Royal Entomological Society of London, 99, 77–128. https://doi.org/10.1111/j.1365-2311.1948.tb01233.x
  166. Kapur, A.P. (1956) Contribution to a knowledge of the fauna of Manipur state, Assam. V. Coleoptera. Records of the Indian Museum, Calcutta, 52, 313–348. https://doi.org/10.26515/rzsi/v52/i2-4/1955/162043
  167. Kapur, A.P. (1957) Variation in the colour pattern of certain ladybird beetles from high altitudes in the Himalayas. Bulletin of the National Institute of Sciences of India, 9, 269–273.
  168. Kapur, A.P. (1958) Coccinellidae of Nepal. Records of the Indian Museum, 53, 309–338.
  169. Kapur, A.P. (1961) Some new or a little known species of Coccinellidae (Insecta: Coleoptera). Part 1. Four new species of Epilachninae from India and Burma. Records of the Indian Museum, 59, 131–141. https://doi.org/10.26515/rzsi/v59/i1-2/1961/161577
  170. Kapur, A.P. (1962) Geographical variations in the colour pattterns of some Indian Ladybeetles (Coccinellidae: Coleoptera). Part I. Coccinella septempunctata Linn., C. transversalis Fabr., and Coelophora bissellata Muls. Proceedings of the First All India Congress of Zoology, 2, 479–492. [1959]
  171. Kapur, A.P. (1963a) The Coccinellidae of the third Mount Everest expedition, 1924 (Coleoptera). Bulletin of the British Museum (Natural History), Entomology, 14, 1–48. https://doi.org/10.5962/bhl.part.8780
  172. Kapur, A.P. (1963b) The taxonomic status and further description of Harmonia expallida Weise (Col., Coccinellidae), feeding on Adelges species (Hem., Adelgidae) in North–western India. Entomophaga, 3, 199–203. https://doi.org/10.1007/BF02376089
  173. Kapur, A.P. (1963) Confirmation of the occurrence of the lady-beetle, Coelophora pupillata (Swartz) (Coccinellidae, Coleoptera) in India and its biological importance. Science and Culture, 29, 264.
  174. Kapur, A.P. (1964) Zoological results of the Indian Cho-Oyu Expedition in Nepal. Part 9. Insecta. Records of the Indian Museum, 59, 269–303. https://doi.org/10.26515/rzsi/v59/i3/1961/161603
  175. Kapur, A.P. (1967) The Coccinellidae (Coleoptera) of the Andamans. Proceedings of the National Institute of Sciences of India, 32 (B), 148–189. [1966]
  176. Kapur, A.P. (1970) Phylogeny of ladybeetles. Proceedings, 57th Indian Science Congress, Kharagpur, Section of Zoology and Entomology, II, 1–14.
  177. Kapur, A.P. (1972) The Coccinellidae (Coleoptera) of Goa. Records of the Zoological Survey of India, 66, 309–320. https://doi.org/10.26515/rzsi/v66/i1-4/1968/161511
  178. Kapur, A.P. (1973) The Coccinellidae (Coleoptera) of the Italian expedition to Karakoram and Hindu Kush. Records of the Zoological Survey of India, 67, 373–378.
  179. Kapur, A.P. & Bhaumik, A.R. (1966) A note on the lady-bird beetles (Coleoptera: Coccinellidae) from Rajasthan with first record of Bulaea lichatschovi (Hummel) from India. Records of the Indian Museum, 59, 445–448. https://doi.org/10.26515/rzsi/v59/i4/1961/161590
  180. Kapur, A.P. & Sudha Rao, V. (1962) A new species of ladybird beetle (Col., Coccinellidae) feeding on Adelges spp. (Hemiptera: Adelgidae) in North western India. Entomophaga, 7, 47–51. https://doi.org/10.1007/BF02375991
  181. Khan, M.A. & Nighat (1991) Insect pest complex of mulberry (Morus spp.) in Kashmir. Journal of Tree Sciences, 10 (1), 8–10.
  182. Khan, I., Din, S., Khalil, S.K. & Rafi, M.A. (2007) Survey of predatory coccinellids (Coleoptera: Coccinellidae) in the Chitral District, Pakistan, Journal of Insect Science, 7 (1), 7. https://doi.org/10.1673/031.007.0701
  183. Kirby, W. & Spence, W. (1828) An Introduction to Entomology: or, Elements of the Natural History of Insects. Vol. III. 5th Edition. Longman, Rees, Orme, Brown & Green, London, 732 pp. https://doi.org/10.5962/bhl.title.152792
  184. Koch, R.L. (2003) The multicolored Asian lady beetle, Harmonia axyridis: A review of its biology, uses in biological control, and non–target impacts. Journal of Insect Science, 3 (32), 1–16.[https://academic.oup.com/insect-science] https://doi.org/10.1673/031.003.3201
  185. Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp.
  186. Korschefsky, R. (1933) Entomological investigations on the spike disease of sandal (16). Coccinellidae (Col.). Indian Forest Records, 19 (6), 1–9.
  187. Korschefsky, R. (1935) Coccinellidae. In: Visser, P.C. & Visser, J.H. (Eds.), Wissenschaftliche ergebnisse der Niederländischen Expeditionen in den Karakorum und die angrenzenden gebiete in den jahren 1922, 1925 und 1929/30. Vol. 1. F.A. Brockhaus, Leipzig, pp. 299
  188. Korschefsky, R. (1940) Vier neue Coccinelliden der Alten Welt. Entomologische Blätter für Biologie und Systematik der Käfer, 36, 1–3.
  189. Korschefsky, R. (1943) Ueber Coccinelliden von den Neuen Hebriden aus dem National-Museum, Paris. Entomologische Blätter, 39, 51–52.
  190. Kovář, I. (2005) Revision of the Palaearctic species of the Coccinella transversoguttata species group with notes on some other species of the genus (Coleoptera: Coccinellidae). Acta Entomologica Musei Nationalis Pragae, 45, 129–164.
  191. Kovář, I. (2007) Family Coccinellidae Latreille, 1807. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 568–631.
  192. Krishnaswamy, S. & Chacko, M.J. (1988) Population dynamics of Adelges knucheli and its predators in the north-western Himalayas I. Host phase: Gallicola. Journal of Biological Control, 2 (2), 92–96.
  193. Kumar, R., Mittal, V., Patankar, N.V. & Ramamurthy, V.V. (2010) Bionomics of mycophagous coccinellid, Psyllobora bisoctonotata (Mulsant) (Coleoptera: Coccinellidae). Munis Entomology & Zoology, 5 (2), 652–657.
  194. Kundoo, A.A. (2019) Taxonomic studies on Coccinellidae in agro-ecosystems of Kashmir and Ladakh. Ph.D. Thesis, Sher-e-Kashmir University of Agricultural Sciences & Technology of Kashmir, Sopore, 87 + xvii pp.
  195. Kurisaki, J. (1914) On two new species of Japanese Coccinellidae. Konchu Sekai, 18, 441–443.
  196. Kurisaki, M. (1923) Descriptions of some interesting coccinellids. Dobutsugaku Zasshi, 35 (413), 101–108. https://doi.org/10.5026/jgeography.35.413
  197. Kuznetsov, V.N. (1977) The biology of the ladybug Aiolocaria mirabilis Motsch. (Coleoptera, Coccinellidae) in Primorye, USSR. Trudy Dal’Nevostochnogo Nauchny Tsentral Biologo-Pochvenny Instituta, 44, 108–117.
  198. Kuznetsov, V.N. (1997) Lady Beetles of the Russian Far East. Memoir No. 1. Centre for Systematic Entomology, The Sandhill Crane Press, Gainesville, Florida, 248 pp.
  199. Kuznetsov, V.N. & Pang, H. (2002) Employment of Chinese Coccinellidae in biological control of aphids in greenhouse in Primorye. Far Eastern Entomologist, 119, 1–5.
  200. Lefroy, H.M. (1909) Indian insect life: a manual of the insects of the plains (tropical India). Agricultural Research Institute, Pusa and Thacker, Spink & Co., Calcutta, 786 pp.
  201. Lewis, G. (1896) On the Coccinellidae of Japan. Annals and Magazine of Natural History, 6, 22–41. https://doi.org/10.1080/00222939608680319
  202. Li, J., Ślipiński, A. & Pang, H. (2010) Revision of the genus Macroilleis (Coleoptera: Coccinellidae: Coccinellini). Annales Zoologici, 60 (1), 15–21. https://doi.org/10.3161/000345410X499470
  203. Linnaeus, C. (1758) Systema Naturae. 10th Edition. L. Salvius, Stockholm, 826 pp.
  204. Liu, C. L. (1950) Studies on the feeding capacity and life history of a mycophagous coccinellid, Halyzia hauseri Mader in Kunming. Annales Entomologici Sinici, 1, 1–15. [in Chinese]
  205. Mader, L. (1926–1934) Evidenz der palaarktischen Coccinelliden nnd ihrer aberrationen in Wort und Bild, l-Teil, Epilachnini, Coccinellini, Halyziini und Synonychini. Zeitschrift des Vereins der Naturbeobachter und Sammler, Wien, 1926 (1), 1927 (2), 1928 (3), 1929 (4), 1930 (5), 1931 (6), 1932 (7), 1933 (8), 1934 (9), 1–24, 25–48, 49–76, 77–124, 124–168, 169–204, 205–244, 245–288, 289–336.
  206. Mader, L. (1930) Neue Coccinelliden aus Yün–nan und Sze–tschwan (China). Entomologischer Anzeiger, 10, 161–166.
  207. Mader, L. (1935–1936) Evidenz der palaarktischen Coccinelliden und ihrer aberrationen in Wort und Bild, l-Teil, Epilachnini, Coccinellini, Halyziini und Synonychini. Entomologischer Anzeiger, 1935 (15) & 1936 (16), 329–372 & 373–412. https://doi.org/10.1086/481877
  208. Mader, L. (1937) Evidenz der paläarktischen Coccinelliden und ihrer Aberrationen in Wort und Bild. Entomologische Nachrichten, Troppau, 11, 41–66.
  209. Magro, A., Lecompte, E., Magne, F., Hemptinne, J.L. & Crouau-Roy, B. (2010) Phylogeny of ladybirds (Coleoptera: Coccinellidae): are the subfamilies monophyletic? Molecular Phylogenetics and Evolution, 54, 833–848. https://doi.org/10.1016/j.ympev.2009.10.022
  210. Majerus, M.E.N. (1994) Ladybirds. Harper Collins, London, 367 pp.
  211. Malhotra, C.P. & Krishnaswami, S. (1962) Coptosoma ostensum Dist.—a pentatomid pest of Palas (Butea monosperma) with notes on its coccinellid predator, Synia melanaria var. rougeti Mulsant and egg parasite, Telenomus species. Indian Forester, 88, 231–237.
  212. Maqbool, A., Ahmed, I., Kieltyk, P. & Ceryngier, P. (2018) Dinocampus coccinellae (Hymenoptera: Braconidae) utilizes both Coccinellini and Chilocorini (Coleoptera: Coccinellidae: Coccinellinae) as hosts in Kashmir Himalayas. European Journal of Entomology, 115, 332–338. https://doi.org/10.14411/eje.2018.033
  213. Maqbool, A., Rather, S.U., Akbar, S.A. & Wachkoo, A.A. (2020) Preliminary survey of ladybird beetle composition (Coleoptera: Coccinellidae) in unmanaged apple orchard ecosystems of Kashmir Himalayas. Proceedings of the Zoological Society, 73, 160–174. https://doi.org/10.1007/s12595-020-00322-w
  214. Mitchell, R.G. & Wright, K.H. (1967) Foreign predator introductions for control of the balsam woolly aphid in the Pacific Northwest. Journal of Economic Entomology, 60, 140–147. https://doi.org/10.1093/jee/60.1.140
  215. Miwa, Y. (1931) A systematic catalogue of Formosan Coleoptera. Report of the Department of Agriculture, Government Research Institute, Taihoku, 55, 1–359.
  216. Miwa, Y. & Yoshida, T. (1935) Catalogue of Japanese Insects. Fasc. IX. Coleoptera. Coccinellidae. (In Japanese). Entomological World, Tokyo, 3 (22), 31–53.
  217. Miyatake, M. (1965) Some Coccinellidae (excluding Scymnini) of Formosa (Coleoptera). Special Bulletin of the Lepidopterists’ Society of Japan, 1, 50–74.
  218. Miyatake, M. (1967) Notes on some Coccinellidae from Nepal and Darjeeling district of India (Coleoptera). Transactions of the Shikoku Entomological Society, 9, 69–78.
  219. Miyatake, M. (1972) A new Formosan species belonging to the genus Singhikalia Kapur, with proposal of a new tribe (Coleoptera: Coccinellidae). Transactions of the Shikoku Entomological Society, 11 (3), 92–98.
  220. Miyatake, M. (1985) Coccinellidae collected by the Hokkaido University Expedition to Nepal Himalaya, 1968 (Coleoptera). Insecta Matsumurana, New Series, 30, 1–33.
  221. Monteith, G.B. (2006) Maternal care in Australian oncomerine shield bugs (Insecta, Heteroptera, Tessaratomidae). Denisia, 19, 1135–1152.
  222. Motschulsky, V. (1860) Insectes du Japon. Etudes Entomologiques, 9, 4–41.
  223. Mulsant, E. (1846) Securipalpes. Vol. 4. Histoire Naturelle des Coleopteres de France, Paris, 380 pp., 1 pl.
  224. Mulsant, E. (1850) Species des Coléoptères Trimères Sécuripalpes. Annales des Sciences Physiques et Naturelles, d’Agriculture et d’Industrie, publiées par la Société nationale d’Agriculture, etc., de Lyon, Deuxième Série, 2, 1–1104. https://doi.org/10.5962/bhl.title.8953
  225. Mulsant, E. (1853a) Supplément à la monographie des coléoptères trimères sécuripalpes. Annales de la Société Linnéenne de Lyon, New Series, 1, 129–333.
  226. Mulsant, E. (1853b) Supplement a la monographie des Coléoptères trimères sécuripalpes. Opuscules Entomologiques, 3, 1–178.
  227. Mulsant, E. (1866) Monographie des Coccinellides. 1re Partie Coccinelliens. Savy et Deyrolle, Paris, 294 pp.
  228. Nagaraja Rao, P.R. & Abraham, E.V. (1959) Occurrence of Alesia discolor Fab. in rice fields in Madras. Madras Agricultural Journal, 46, 234–235.
  229. Nagarkatti, S. & Ghani, M.A. (1972) Ecology and biology of predators. Coleoptera: Coccinellidae. In: Rao, V.P. & Ghani, M.A. (Eds.), Studies on predators of Adelges spp. in the Himalayas . Miscellaneous Publication No. 3. Commonwealth Institute of Biological Control, Indian Statio, pp. 58–88
  230. Nakane, T. & Araki, M. (1959) Entomological results from the scientific survey of the Tokara Islands. VI. Coleoptera: Coccinellidae. Scientific Reports of Kyoto Prefectural University (Natural and Living Science), 3A, 45–52.
  231. Narasimham, A.U. & Sankaran, T. (1985) Some mealybugs and their natural enemies in India. Proceedings of the National Seminar on Entomophagous Insects, Calicut, 1985, 29–36
  232. Nunenmacher, F.W. (1912) Studies amongst the Coccinellidae, No. 4 (Col.). In: Entomological News and Proceedings of the Entomological Section. 23 (10). Academy of Natural Sciences, Philadelphia, pp. 448–451.
  233. Obrycki, J.J. & Kring, T.J. (1998) Predaceous Coccinellidae in biological control. Annual Review of Entomology, 43, 295–321. https://doi.org/10.1146/annurev.ento.43.1.295
  234. Omkar & Pervez, A. (1999) New records of coccinellids from Uttar Pradesh I. Journal of Advanced Zoology, 20 (2), 102–106.
  235. Omkar & Pervez, A. (2004) Predaceous coccinellids in India: predator-prey catalogue. Oriental Insects, 38, 27–61. https://doi.org/10.1080/00305316.2004.10417373
  236. Ozbek, H. & Cetin, G. (1991) Contribution to the fauna of Coccinellidae (Coleoptera) from eastern Anatolia along with some new records from Turkey. Turkish Journal of Entomology, 15 (4), 193–202.
  237. Pallas, P.S. (1773) Reise durch verschiedene Provinzen des Rußischen Reichs. Zweyter Theil. zweytes Buch vom Jahr 1771. Kayserliche Academie der Wißenschaften, St. Petersburg. [unknown pagination]
  238. Pang, X.F. & Mao, J.L. (1979) Coleoptera: Coccinellidae, II. Khongguo Jingi Kunchang Zhi, 14, 1–170.
  239. Pang, H., Huang, B. & Pang, X.-F. (2002) Coleoptera: Coccinellidae. In: Fauna of Insects in Fujian Province of China. Vol. 6. Fujian Science Technology Publishing House, Fuzhou, Fujian, pp. 281–357.
  240. Patankar, N.V., Mittal, V., Kumar, R. & Ramamurthy, V.V. (2009) Can Psyllobora bisoctonotata and Illeis cincta (Coccinellidae: Coleoptera) be advocated for biological control of powdery mildews? Indian Journal of Agricultural Sciences, 79 (6), 488–490.
  241. Phaloura, S.P.S. & Singh, T. (1991) Larval chaetotaxy of three species of genus Harmonia Mulsant (Coleoptera: Coccinellidae) and phylogenetic relationship amongst its species. Elytron, 5, 39–46.
  242. Phaloura, S.P.S. & Singh, T. (1993) Biological observations on three species of aphidophagous coccinellids of genus Harmonia Mulsant. Journal of Insect Science, 6, 246–249.
  243. Phaloura, S.P.S., Singh, T. & Singh, T. (1992) Biological observations on Aiolocaria hexaspilota (Hope) (Coleotera: Coccinellidae)–a predator on the immature stages of a chrysomelid (Coleoptera)—Chrysomela chlorina Maulik. Journal of Ecobiology, 4, 33–37.
  244. Ponnusamy, N., Biwash, G. & Pal, S. (2021) Biology and feeding potential of giant ladybird beetle, Anisolemnia dilatata (Fab.) (Coleoptera: Coccinellidae) on som aphids (Aiceona sp.) (Hemiptera: Aphididae). Pest Management in Horticultural Ecosystems, 20 (1), 61–64. https://doi.org/10.5958/0974-4541.2021.00011.4
  245. Pontoppidan, E. (1763) Den Danske Atlas. Tomus 1, 4°. Godiche, Kiöbenhavn, 669 pp.
  246. Pope, R.D. (1987) The Coccinellidae (Coleoptera) described by C. P. Thunberg. Entomologica Scandinavica, 18 (1), 51–66. https://doi.org/10.1163/187631287X00025
  247. Pope, R.D. (1989) A revision of the Australian Coccinellidae (Coleoptera). Part I. Subfamily Coccinellinae. Invertebrate Taxonomy, 3, 633–735. https://doi.org/10.1071/IT9880633
  248. Poorani, J. (2002a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307–383. https://doi.org/10.1080/00305316.2002.10417335
  249. Poorani, J. (2002b) A review of the genus Oenopia Mulsant (Coleoptera: Coccinellidae) of the Indian subcontinent, with description of a new species. Oriental Insects, 36, 97–116. https://doi.org/10.1080/00305316.2002.10417328
  250. Poorani, J. (2005) Notes on the Coccinellidae (Coleoptera) of the Indian subcontinent, including new synonymies. Journal of Biological Control, 18 (2), 185–187.
  251. Poorani, J. (2007) First record of Hippodamia variegata (Goeze) (Coleoptera: Coccinellidae) from South India. Journal of Biological Control, 21 (2), 295–296.
  252. Poorani, J. (2014) Calvia explanata sp. n. (Coleoptera: Coccinellidae: Coccinellinae) from the Indian Subcontinent. Biodiversity Data Journal, 2, e1164. https://doi.org/10.3897/BDJ.2.e1164
  253. Poorani, J. (2014) An unusual new species of Micraspis Chevrolat (Coleoptera: Coccinellidae) from northeastern India. Biodiversity Data Journal, 2, e4112, https://doi.org/10.3897/BDJ.2.e4112
  254. Poorani, J. (2019) A new species of Micraspis Chevrolat (Coleoptera: Coccinellidae) from Arunachal Pradesh, India. Zootaxa, 4559 (1), 189–192. https://doi.org/10.11646/zootaxa.4559.1.11
  255. Poorani, J. & Booth, R.G. (2005) On the identity of Dysis excellens Crotch (Coleoptera: Coccinellidae: Coccinellini), a little known Oriental lady beetle. Zootaxa, 875 (1), 1–5. https://doi.org/10.11646/zootaxa.875.1.1
  256. Poorani, J. & Booth, R.G. (2006) A new sibling species of Halyzia straminea (Hope) (Coleoptera: Coccinellidae: Coccinellinae) from the Indian subcontinent. Zootaxa, 1354 (1), 63–68. https://doi.org/10.11646/zootaxa.1354.1.5
  257. Poorani, J. & Ślipiński, A. (2005) Notes on the genus Protothea Weise with redescription of P. quadripunctata (Mulsant) (Coleoptera: Coccinellidae: Coccinellini). Annales Zoologici, 55 (3), 329–333.
  258. Poorani, J. & Thangjam, R. (2019) New additions to Indian fauna of Coccinellidae (Coleoptera). Oriental Insects, 53 (4), 547–565. https://doi.org/10.1080/00305316.2018.1550448
  259. Poorani, J., Ślipiński, A. & Booth, R.G. (2008) A revision of Synona Pope (Coleoptera: Coccinellidae: Coccinellini). Annales Zoologici, 58, 579–594. https://doi.org/10.3161/000345408X364427
  260. Poorani, J., Jalali, S.K. & Ojha, R. (2015) Morphological characters and DNA barcodes to separate Oenopia sauzeti Mulsant and O. mimica Weise (Coleoptera: Coccinellidae), two externally similar lady beetles from the Indian Subcontinent. Entomon, 40 (4), 235–242.
  261. Poorani, J., Sankararaman, H. & Sathyaseelan, V. (2017) First report of Bulaea lividula bocandei Mulsant (Coleoptera: Coccinellidae) from India with notes on its diagnosis, host plants and distribution. Journal of Biological Control, 31 (4), 194–197. https://doi.org/10.18311/jbc/2017/17974
  262. Poorani, J., Sankararaman, H. & Anusree, S.S. (2021) A review of Indian species of Phrynocaria Timberlake (Coleoptera: Coccinellidae) with notes on synonymy and a new species from South India, hitherto misidentified as Coelophora circumusta (Mulsant). Zootaxa, 4926 (1), 117–133. https://doi.org/10.11646/zootaxa.4926.1.8
  263. Poorani, J., Booth, R.G., Anuradha, C., Gracy, R. G., Thanigairaj, R. & Swathi, R.S. (2023) Identity of the ‘true’ Micraspis discolor (Fabricius) (Coleoptera, Coccinellidae) with illustrated diagnostic notes on other Micraspis spp. in Indian paddy fields. Zootaxa, 5271 (3), 446–476. https://doi.org/10.11646/zootaxa.5271.3.2
  264. Puttarudriah, M. & Channabasavanna, G.P. (1952) Two giant ladybird beetles predaceous on aphids in Mysore. Indian Journal of Entomology, 14, 20.
  265. Puttarudriah, M. & Channabasavanna, G.P. (1953) Beneficial coccinellids of Mysore–I. Indian Journal of Entomology, 15, 87–96.
  266. Puttarudriah, M. & Channabasavanna, G.P. (1955) Beneficial coccinellids of Mysore–II. Indian Journal of Entomology, 17, 1–5.
  267. Puttarudriah, M. & Channabasavanna, G.P. (1956) Some beneficial coccinellids of Mysore. Journal of the Bombay Natural History Society, 54, 156–159.
  268. Puttarudriah, M. & Channabasavanna, G.P. (1957) Notes on some predators of mealybugs (Coccidae, Hemiptera). The Mysore Agricultural Journal, 32, 4–19.
  269. Puttarudriah, M. & Maheswari, B.M. (1966) Biology, fecundity and rate of feeding of Synonycha grandis Thunberg. Proceedings, Indian Academy of Sciences, 12B, 265–278.
  270. Rachappa, V., Hegde, R., Kulkarni, K.A. & Patil, R.K. (2002) Biology and feeding potential of Synia melanaria Mulsant on Coptosoma ostensum Distant. Myforest, 38 (2), 145–150.
  271. Rafi, M.A., Irshad, M. & Inayatullah, M. (2005) Predatory ladybird beetles of Pakistan. National Agricultural Research Centre, Islamabad, 105 pp.
  272. Rahman, K.A. (1940) Important insect predators of India. Proceedings of the Indian Academy of Sciences (B), 12, 67–74. https://doi.org/10.1007/BF03049103
  273. Rahman, K.A. & Khan, A.W. (1941) Observations on Aphelinus mali (Hald.) in the Punjab. Indian Journal of Agricultural Sciences, 11, 446–450.
  274. Rahman, M.J., Ali, M.S. & Mannan, M.A. (1993). Illeis indica Timberlake (Coleoptera: Coccinelldae), a mildew feeder in mulberry. Bulletin of Sericultural Research, 4, 80–81.
  275. Raimundo, A.A.C. & Harten, A. van (2000) An annotated checklist of the Coccinellidae (Insecta: Coleoptera) of Yemen. Fauna of Arabia, 18, 211–243.
  276. Raimundo, A.A.C., Fürsch, H. & Harten, A. van (2008) Order Coleoptera, family Coccinellidae. Arthropod Fauna of UAE, 1, 217–239.
  277. Ramani, S., Poorani, J. & Bhumannavar, B.S. (2002) Spiralling whitefly, Aleurodicus dispersus, in India. Biocontrol News and Information, 23 (2), 55–62.
  278. Ramaraju, K. & Poorani, J. (2012) A new species of Coccipolipus (Acari: Podapolipidae) parasitic on the giant coccinellid beetle from India. International Journal of Acarology, 38, 260–296. https://doi.org/10.1080/01647954.2011.651156
  279. Ramírez, J., González, G. & Sánchez, Y. (2018) First record of Cheilomenes sexmaculata (Fabricius, 1781) (Coleoptera, Coccinellidae) from Colombia. Check List, 14 (1), 77–80. https://doi.org/10.15560/14.1.77
  280. Rao, V.P. (1969) Survey for natural enemies of aphids in India. CIBC-US PL-480 Project, Final Technical Report. Report of the Commonwealth Institute of Biological Control, 480, 1–93.
  281. Rao, V.P. (1969) India as a source of natural enemies of pests of citrus. Proceedings, First International Citrus Symposium, 2, 785–792.
  282. Rao, V.P. & So, P.Y. (1967) Notes on Coccinellidae of Hong Kong. Technical Bulletin, Commonwealth Institute of Biological Control, 9, 9–24.
  283. Rao, V.P., Datta, B. & Ramaseshaiah, G. (1970) Natural enemy complex of flushworm and phytophagous mites on tea in India. Scientific Publication Series. No. 5. Tea Board, India, Calcutta, 53 pp.
  284. Rao, V.P., Ghani, M.A., Sankaran, T. & Mathur, K.C. (1971) A review of the biological control of insects and other pests in South-East Asia and the Pacific region. Technical Communication, Commonwealth Institute of Biological Control, Slough, 149 pp.
  285. Rawat, R.R. & Modi, B.N. (1969b) Record of some predacious beetles on coccid, aphid and mite pests from Madhya Pradesh. Indian Journal of Agricultural Sciences, 39, 1057–1060.
  286. Reiche, M. L. (1861) Notes synonymiques. Annales de la Société entomologique de France, Serie 4, 1, 211–1212.
  287. Ren, S. X., Wang, X. M., Pang, H., Peng, Z. Q. & Zeng, T. (2009) Colored pictorial handbook of ladybird beetles in China. Science Press, Beijing, 336 pp. [in Chinese]
  288. Rhamhalinghan, M. (1989) Studies on Coelophora bissellata Muls. (Coleoptera: Coccinellidae). 1. Field recognition of different larval instars and the traits of pupae and adults. Journal of the Bombay Natural History Society, 86, 114–117.
  289. Richerson, J.V. (1970) A world list of parasites of Coccinellidae. Journal of the Entomological Society of British Columbia, 67, 33–48.
  290. Roy, P. (1976) Life history and feeding behaviour of Coccinella transversalis Fab. (Coccinellidae, Coleoptera), a predator of mustard aphid, Lipaphis erysimi (Kalt.). Indian Biologist, 8, 59–61.
  291. Rybakow, G. (1889) Insecta in itinere d. N. Przewalski in Asia Centrali Novissimehecta. XIV. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 286–290.
  292. Rye, E.C. (1873) Insecta. In: Zoological Record 1871. Vol. 8. John van Voorst, London. [unknown pagination]
  293. Sajan, K.C., Kapil, K. & Anjali, K. (2018) Records of ladybeetles (Coleoptera: Coccinellidae) from hilly regions of Nepal. Indian Journal of Entomology, 80 (4), 1236–1248. https://doi.org/10.5958/0974-8172.2018.00242.0
  294. Sajan K.C., Větrovec, J. & Kafle, K. (2019) Lady beetles of Nepal (Coleoptera: Coccinellidae): Coccinellinae from the fields at Nepal Agricultural Research Council, Khumaltar, Lalitpur. International Journal of Entomology Research, 4 (4), 157–165.
  295. Sasaji, H. (1968) Coccinellidae collected in the paddy fields of the Orient, with descriptions of new species (Coleoptera). Mushi, 42, 119–132.
  296. Sasaji, H. (1971) Fauna Japonica. Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp.
  297. Sasaji, H. (1977) Larval characters of Asian species of the genus Harmonia Mulsant. Memoirs of the Faculty of Education, Fukui University Series II, 27, 1–17.
  298. Sasaji, H. (1981) Biosystematics of Harmonia axyridis complex (Coleoptera: Coccinellidae). Memoirs of the Faculty of Education, Fukui University Series II, 30, 59–79.
  299. Sasaji, H. (1982) A revision of the Formosan Coccinellidae (III), subfamily Coccinellinae (Coleoptera). Memoirs of the Faculty of Education, Fukui University, Series II, New Series, 31, 1–49.
  300. Sasaji, H. (1992) Descriptions of four coccinellid larvae of Formosa, with the phylogenetic importance (Coleoptera: Coccinellidae). Memoir of the Faculty of Education, Fukui University Series II Natural Science, 42 (1), 1–11.
  301. Sasaji, H. & Akamatsu, M. (1979) Reproductive continuity and genetic relationships in the forms of the genus Menochilus (Coleoptera: Coccinellidae). Memoirs of the Faculty of Education, Fukui University, Series II (Natural Sciences), 29, 1–18.
  302. Sasaji, K. & Tsubokawa, K. (1982) Supplementary descriptions of the coccinellid larvae of Japan (Coleoptera). Memoirs of the Faculty of Education, Fukui University Series II, 32, 33–66.
  303. Sasaji, H., Yahara, R. & Saito, M. (1975) Reproductive isolation and species specificity in two ladybirds of the genus Propylea (Coleoptera). Memoirs of the Faculty of Education, Fukui University, Series II (Natural Sciences), 25, 13–34.
  304. Savoiskaya, G.I. (1970) Coccinellids of the Alma-Ata reserve. Trudy Alma-Atinskogo Gosudarstvennyi Zapovednik, 9, 163–187.
  305. Savoiskaya, G. I. (1983) Larvae of the coccinellid (Coleoptera: Coccinellidae) fauna of the USSR. Nauka, Leningrad, 242 pp. [in Russian]
  306. Savoiskaya, G.I. & Klausnitzer, B. (1973) Morphology and taxonomy of the larvae with keys for their identification. In: Hodek, I. (Ed.), Biology of the Coccinellidae. W. Junk, Prague and Hague, pp. 36–55. https://doi.org/10.1007/978-94-010-2712-0_2
  307. Schaefer, P.W. & Semyanov, V.P. (1992) Arthropod parasites of Coccinella septempunctata (Coleoptera: Coccinellidae): world parasite list and bibliography. Entomological News, 103, 125–134.
  308. Schilder, F.A. & Schilder, M. (1928) Die Nahrung der Coccinelliden und ihre Beziehung zur Verwandschaft der Arten. Arbeiten aus der Biologischen Reichsanstalt für Land–und Forstwirtschaft, 16, 213–282.
  309. Schneider, D.H. (1792) Neustes Magazin fur die Liebhaber der Entomologie. Vol. 2. Stralsund und Leipzig, Struck, pp. 128–256.
  310. Schönherr, C.J. (1808) Synonyma Insectorum, oder: Versuch einer Synonymie aller bisher bekannten Insecten; nach Fabricii Systema Eleutheratorum &c. geordnet. Erster Band. Eleutherata oder Käfer. Zweiter Theil. CF Marquard, Stockholm. [unknown pagination]
  311. Seago, A.E., Giorgi, J.A., Li, J.H. & Ślipiński, A. (2011) Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data. Molecular Phylogenetics and Evolution, 60, 137–151. https://doi.org/10.1016/j.ympev.2011.03.015
  312. Semenow, A. (1895) Coleoptera Asiatica nova. III. Horae Societatis Entomologicae Rossicae, 29, 125–144.
  313. Semyanov, V.P. (1980) Ways to employ the Coccinellidae. Zashchita Rastenii, 8, 20–21. [in Russian]
  314. Semyanov, V.P. (1996) The method of breeding and longtime storage of tropical ladybird species Leis dimidiata (Fabr.) (Coleoptera: Coccinellidae). Entomologicheskoe Obozrenie, 75 (3), 714–720. [in Russian]
  315. Semyanov, V.P. (1999) Biology of ladybirds (Coleoptera, Coccinellidae) from South-East Asia. I. Leis dimidiata (Fabr.). Entomologicheskoe Obozrenie, 78 (3), 537–544. [in Russian].
  316. Semyanov, V.P. (2000) The biology of the coccinellids (Coleoptera: Coccinellidae) from Southeast Asia. II. Harmonia sedecimnotata (F.). Entomologicheskoe Obozrenie, 79, 3–9. [in Russian]
  317. Semyanov, V.P. (2001) The biology of coccinellids (Coleoptera: Coccinellidae) from South–East Asia. III. Lemnia biplagiata (Swartz). Entomologicheskoe Obozrenie, 80, 578–584. [in Russian]
  318. Shantibala, K. & Singh, T.K. (1991) Studies on the vertical distribution of aphidophagous coccinellids (Coleoptera: Coccinellidae) of Manipur and Nagaland. Journal of Aphidology, 5, 39–43.
  319. Sharma, P.K. & Verma, A.K. (1993) Studies on the coccinellid predators of the cabbage aphid Brevicoryne brassicae in Himachal Pradesh. Journal of Biological Control, 7 (2), 15–19.
  320. Sharma, P.L., Verma, S.C., Chandel, R.S., Chandel, R.P.S. & Thakur, P. (2017) An inventory of the predatory Coccinellidae of Himachal Pradesh, India. Journal of Entomology and Zoology Studies, 5 (6), 2503–2507.
  321. Sharmila, M., Lukeshwari, R.K., Singh, T.K. & Agarwala, B.K. (2009) Effect of different prey densities on the reproductive attributes of Harmonia dimidiata (Fabricius) (Coleoptera: Coccinellidae). Entomon, 34 (1), 23–27.
  322. Schönherr, C.J. (1808) Synonymia Insectorum, oder Versuch einer Synonymie Aller bischer bekkanten Insecten nach Fabricii Systema Eleutheratorum etc. geordnet. Erster Band. Eleutherata oder Käfer. Zweuter Theil. Spercheus-Cryptocephalus. Carl Friedr. Marquard, Stockholm, 424 pp.
  323. Sicard, A. (1907) Notes sur quelques Coccinellides Paléarctiques (Col.). Bulletin de la Société Entomologique de France, 1907, 132–136. https://doi.org/10.5962/bhl.part.4570
  324. Sicard, A. (1910) Espèces et variétés nouvelles des Coccinellides de l’Inde (Coleoptera) appartenant à la collection de M. Andrewes (de Londres). Annales de la Société Entomologique de France, 79, 377–389. [1910]
  325. Sicard, A. (1912) Description d’especes et varietes nouvelles de Coccinellides de la collection de Deutsche Entomologisches Museum de Berlin–Dahlem. Archiv für Naturgeschichte, 78 (A6), 129–138.
  326. Sicard, A. (1913) Notes sur quelques Coccinellides de l’Inde et de Birmanie appartenant à la collection de M. Andrewes, de Londres et description d’espèces et de variétés nouvelles. Annales de la Société Entomologique de France, 81, 495–506. [1912]
  327. Singh, K.C. & Singh, T.K. (1985) Aphidophagous coccinellids of north eastern India: Manipur—I. Entomon, 10, 291–295.
  328. Singh, K.C. & Singh, T.K. (1988) The Coccinellidae (Coleoptera) of Mizoram. Bulletin of Entomology, 29 (2), 186–189.
  329. Singh, T. & Phaloura, S.P.S. (1990) A field key to the larvae of four species of genus Harmonia Mulsant (Coleoptera: Coccinellidae). Journal of Entomological Research, 14, 89–90.
  330. Singh, L.S., Singh, K.C. & Singh, T.K. (1993) Biology and feeding potential of Oenopia quadripunctata Kapur, a coccinellid predator of an oak aphid, Tuberculatus (Acanthocallis) nervatus Chakrabarti and Raychaudhuri. Journal of Advanced Zoology, 14, 7–11.
  331. Ślipiński, A. (2007) Australian ladybird beetles (Coleoptera: Coccinellidae). Their biology and classification. Australian Biological Resources Study, Canberra, xviii + 286 pp.
  332. Ślipiński, A., Li, J. & Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https://doi.org/10.1071/9781486303885
  333. Ślipiński, S.A. & Pakaluk, J. (1991) Problems in the classification of the Cerylonid series of Cucujoidea (Coleoptera). In: Zunino, M., Belles, X. & Blas, M. (Eds.), Advances in Coleopterology . European Association of Coleopterology, Barcelona, pp. 79–88.
  334. Ślipiński, A. & Tomaszewska, W. (2010) Coccinellidae Latreille, 1802. In: Leschen, R.A.B., Beutel, R.G. & Lawrence, J.F. (Eds.), Handbook of Zoology. Vol. Coleoptera. Walter de Gruyter GmbH & Co. KG, Berlin and New York, pp. 454–472. https://doi.org/10.1515/9783110911213.454
  335. Stebbing, E.P. (1903) Coleoptera 2. Notes upon the known predaceous Coccinellidae of the Indian region, Part I. Indian Museum Notes, VI (1), 47–62.
  336. Stebbing, E.P. (1914) Indian forest insects of economic importance. Coleoptera. Eyre and Spottiswoode, Ltd., London, 648 pp. https://doi.org/10.5962/bhl.title.9203
  337. Subramaniam, T.V. (1924a) A note on colour variations in a common ladybird beetle, Chilomenes sexmaculata Fb. In: Fletcher, T.B. (Ed.), Report of the Proceedings of the Fifth Entomological Meeting, Pusa, 1924. [unknown pagination]
  338. Subramaniam, T.V. (1924b) Some coccinellids of South India. In: Fletcher, T.B. (Ed.), Report of the Proceedings of the Fifth Entomological Meeting, 1923, pp. 108–118
  339. Subramaniam, V.K. (1954–1955) Control of the fluted scale in peninsular India. Indian Journal of Entomology, 16 & 17, 391–415 & 103–120.
  340. Subramanyam, T.V. (1925) Coptosoma ostensum Dist. and its enemy Synia melanaria Muls. Journal of the Bombay Natural History Society, 30, 924–925.
  341. Sudha Rao, V. (1962) The status of Coccinella septempunctata L. and its varieties divaricata Oliv. and confusa Wied. Canadian Entomologist, 94, 1341–1343. https://doi.org/10.4039/Ent941341-12
  342. Thompson, W.R. & Simmonds, F.J. (1964) A Catalogue of the Parasites and Predators of Insect Pests. Section 3. Predator-Host Catalogue. Commonwealth Agricultural Bureaux, Farnham Royal, 208 pp.
  343. Thunberg, C.P. (1781) Dissertatio Entomologica Novas Insectorum Species, sistens cujus partem primam, Cons. Exper. Facul. Med. Upsal., publice ventilandam exhibent praeses Carol. P. Thunberg, et respondens Samuel Nicol. Casström. Joh. Edman, Upsaliae, 28 pp. https://doi.org/10.5962/bhl.title.12285
  344. Thunberg, C.P. (1820) Coleoptera Capensis Antennarum clava solida et perfoliata. Memoires de l’Academie Imperiale des Sciences de Saint Petersbourg, 7, 362–372.
  345. Timberlake, P.H. (1943) The Coccinellidae or lady beetles of the Koebele collection–Part I. Hawaiian Planters’ Record, 47, 1–67.
  346. Tomaszewska, W., Escalona, H.E., Hartley, D., Li, J., Wang, X., Li, H.S., Pang, H., Ślipiński, A. & Zwick, A. (2021) Phylogeny of true ladybird beetles (Coccinellidae: Coccinellini) reveals pervasive convergent evolution and a rapid Cenozoic radiation. Systematic Entomology, 46, 611–631. https://doi.org/10.1111/syen.12479
  347. Ukrainsky, A.S. (2006) Five new replacement ladybird (Coleoptera: Coccinellidae) generic names. Russian Entomological Journal, 15 (4), 399–400.
  348. Vandenberg, N.J. (1996) Notes on Oenopia Mulsant (Coleoptera: Coccinellidae). Proceedings of the Entomological Society of Washington, 98 (3), 385–389.
  349. Vandenberg, N.J. (2002a) Family 93. Coccinellidae Latreille 1807. In: Arnett, R.H., Thomas, Jr. M.C., Skelley, P.E. & Frank, J.H. (Eds.), American Beetles. Vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, Florida, pp. 371–389.
  350. Vandenberg, N.J. (2002b) The new world genus Cycloneda Crotch (Coleoptera: Coccinellidae: Coccinellini): Historical review, new diagnosis, new generic and specific synonyms, and an improved key to North American species. Proceedings of the Entomological Society of Washington, 104, 221–236.
  351. Vandenberg, N.J. & Gordon, R.D. (1991) Farewell to Pania Mulsant (Coleoptera: Coccinellidae): a new synonym of Propylea Mulsant. Coccinella, 3, 30–35.
  352. Veenakumari, K. & Mohanraj, P. (2007) Additions to the coccinellid fauna of the Andaman Islands and the biology of the endemic Chilocorus coelosimilis Kapur 1966 (Coleoptera: Coccinellidae). Journal of the Bombay Natural History Society, 104 (2), 232–235.
  353. Wadhi, S.R. & Parshad, B. (1980) Some new records of Coccinellidae from Nepal Himalayas. Bulletin of Entomology, 21, 144–147.
  354. Walker, F. (1859) Characters of some apparently undescribed Ceylon insects. Annals and Magazine of Natural History, Series 3, 4, 217–224. https://doi.org/10.1080/00222935908697111
  355. Watson, W.Y. (1956) A study of the phylogeny of the genera of the tribe Coccinellini (Coleoptera). Contributions of the Royal Ontario Museum, Division of Zoology and Palaentology, 42, 1–52. https://doi.org/10.5962/bhl.title.52186
  356. Wiedemann, C.R.G. (1823) Zweihundert neue Käfer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin, 2 (1), 3–133.
  357. Weise, J. (1879) Beiträge zur Käfer fauna von Japan. Deutsche Entomologische Zeitschrift, 23, 149–152. https://doi.org/10.1002/mmnd.48018790119
  358. Weise, J. (1885) Bestimmungs-Tabellen der europäischen Coleopteren. II. Heft. Coccinellidae. II. Auflage mit Berücksichtigung der Arten aus dem nöordlichen Asien, Modling, 83 pp.
  359. Weise, J. (1889) Insecta a Cl. G. N. Potanin in China et in Mongolia novissime lecta. IX. Chrysomelidae et Coccinellidae. Horae Societatis Entomologicae Rossicae, 23, 560–653.
  360. Weise, J. (1891) Neue Coccinelliden. Deutsche Entomologische Zeitschrift, 35, 282–288. https://doi.org/10.1002/mmnd.48018910212
  361. Weise, J. (1892) Les Coccinellides du Chota–Nagpore. Annales de la Société Entomologique du Belgique, 36, 16–30.
  362. Weise, J. (1895a) Neue Coccinelliden sowie Bemerkungen zu bekannten Arten. Annales de la Société Entomologique du Belgique, 1895, 120–146.
  363. Weise, J. (1895b) Insectes du Bengale. Coccinellidae. Annales de la Société Entomologique du Belgique, 1895, 151–157.
  364. Weise, J. (1897) Coccinellen aus Ostafrika (Usambara). Deutsche Entomologische Zeitschrift, 41, 289–304. https://doi.org/10.1002/mmnd.48018970120
  365. Weise, J. (1898) Ueber bekannte und neue Coccinelliden. Archiv für Naturgeschichte, Band I, 64, 225–238.
  366. Weise, J. (1900) Coccinelliden aus Ceylon gesammelt von Dr. Horn. Deutsche Entomologische Zeitschrift, 44, 417–448. https://doi.org/10.1002/mmnd.48019000237
  367. Weise, J. (1901) Contribution a l’etude de la faune entomologique de Sumatra (Chasses de M.J.-L.Weyers) xve article. Coccinellides. Annales de la Société Entomologique du Belgique, 45, 91–96.
  368. Weise, J. (1902) Coccinelliden aus der Sammlung des ungarischen National Museums. Természetrajzi Füzetek, 25, 489–520.
  369. Weise, J. (1905) Ueber Coccinelliden. Deutsche Entomologische Zeitschrift, 49, 217–220. https://doi.org/10.1002/mmnd.48119050313
  370. Weise, J. (1907) Kleine Mitteilungen und synonymische Bemerkungen. Entomologische Zeitung, Wien, 26, 13–15. https://doi.org/10.5962/bhl.part.8861
  371. Weise, J. (1908) Coleopteren aus Ostindien. Stettiner Entomologische Zeitung, 69, 213–230.
  372. Weise, J. (1910) Chrysomeliden und Coccinelliden. Brünn Verhandlungen des Naturforschenden Vereines, 48 (1909), 25–53.
  373. Weise, J. (1912) Über Hispien und Coccinelliden. Archiv für Naturgeschichte, 78, 101–120.
  374. Weise, J. (1923) H. Sauter’s Formosan-Ausbeute: Coccinelliden. Archiv für Naturgeschicht, 89 (A), 2, 182–189.
  375. Xiao, N. & Li, H. (1992) Coleoptera: Coccinellidae. In: Huang, F. (Eds.), Insects of Wuling Mountains area, southwestern China. Science Press, Beijing, pp. 368–390. [in Chinese with English summary]
  376. Yablokov-Khnzoryan, S.M. (1977) A new species of ladybeetle from India (Coleoptera: Coccinellidae). Doklady Akademii Nauk Armyanskoi SSR, 64 (1), 61–63.
  377. Yablokov-Khnzoryan, S. M. (1978) Two new species of ladybeetles from Eastern Asia (Coleoptera, Coccinellidae). Doklady Akademii Nauk Armyanskoi SSR, 67 (3), 180–183.
  378. Yu, G. (1994) Cladistic analyses of the Coccinellidae (Coleoptera). Entomologia Sinica, 1 (1), 17–30. https://doi.org/10.1111/j.1744-7917.1994.tb00191.x
  379. Yu, G. (1998) Notes on the genus Alloneda Iablokoff-Khnzorian with description of a new species (Coleoptera: Coccinellidae). Zoological Studies, 37, 282–285.
  380. Yu, G. (2010) Chinese Lady Beetles (The Subfamily Coccinellinae). China Scientific Press, Beijing, 180 pp. [in Chinese]
  381. Yu, G. (2011) The Coccinellidae of Taiwan. Chemical Industry Press, Beijing, 198 pp. [in Chinese with English summary]
  382. Yu, G. & Wang, H.Y. (1999) Guidebook to lady beetles of Taiwan. Shih Pui Ni, Taipei, 230 pp. [bilingual, in English and Chinese]
  383. Zimsen, E. (1964) The type material of J.C. Fabricius. Munksgaard, Copenhagen, 656 pp.